单项选择题

下列关于真核细胞与原核细胞的比较叙述错误的是（）

A.真核细胞分化出细胞核和细胞质，形成了结构和功能特异的各种膜相细胞器
B.真核细胞有核骨架和细胞质骨架
C.真核细胞的染色体DNA是环状的，但与组蛋白和非组蛋白结合形成复杂的结构
D.真核细胞基因表达更复杂，具有多层次性

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